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- Adaptive management - Monitoring
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Adaptive management
It has been emphasised throughout this study that the population
dynamics of hippo are most unusual and the population response
to management interventions is difficult to predict. What
has been put forward as the expected population dynamics of
hippo in response to various management treatments should
be seen as hypotheses in an adaptive management system (Holling
1976). Indeed, the proposed 'project' which involves harvesting,
sport hunting and problem animal control should be seen as
an active adaptive management research opportunity.
Under Passive Adaptive Management ("cautious fiddling") the
effort is varied narrowly around some point for fear that
any major change will result in system collapse. The result
is that the optimum operating point is seldom detected. Under
Active Adaptive Management the effort is deliberately varied
over a period of time so that the population characteristic
(or response to the harvest) can be defined. This may have
considerable relevance to the hippo 'project'.
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Monitoring
To understand the population dynamics of the Caprivi hippo
several key parameters need to be monitored:
Population numbers: Once population harvesting starts,
an aerial survey such as that done last year by Stander (2004)
should be carried out annually. To make comparisons from one
year to the next it is worth maintaining the same strata and
counting blocks. The ecological objective of this management
plan is to create the conditions for the hippo population
to increase and such surveys will be needed to establish whether
this is happening.
This should be supplemented with local level counts on the
ground (or in the water) in all the conservancies. Hippo are
one of the few species whose numbers can be reasonably measured
by direct observation. If conservancies are to be seen to
be genuinely managing their wildlife then this sort of inventory
is necessary and it should not be confused with Event Book
monitoring. It requires one or two individuals to spend long
hours with binoculars getting to know exactly what hippo are
in their areas. The numbers to keep track of are fairly low:
Salambala - 1; Kwandu - 8; Mashi - 18; Mayuni - 42; Kasika
and Impalila -30). This data is useful also for 'ground-truthing'
aerial surveys.
This intensive ground observation is also an excellent method
of monitoring illegal hunting. Apart from the likelihood of
actually seeing illegal hunters, the disappearance of known
animals from local populations is a good indicator that illegal
hunting is taking place.
Ages of all hippo killed: All lower jaws from hippo
killed or dying naturally should be collected, clearly tagged:
Each lower mandible should carry a label giving date of death,
locality and category of death (e.g. natural mortality, hunting
trophy, problem animal or animal harvested). The jaws should
be transported to a central place where one person can determine
the ages of
the animals. Over several years an age structure for the
population can be built up and, in conjunction with the population
model of this study, a great deal can be inferred about the
population dynamics.
This is also the most reliable method for adjusting quotas
of trophy animals: the hypothesis put forward in this study
is that a half-percent quota will result in a few males living
beyond 30 years of age. If this is not the case, then the
quota should be reduced: alternatively, if most trophies are
over 30 years of age, quotas can be increased. These data
also allow some inference about the age-specific natural mortality
for the older males.
Reproductive data: For all females killed under a
harvesting programme, it is worth recording whether or not
they are pregnant or lactating in order to establish both
fecundity and age at first conception for the population since
these reproductive
parameters are variable in the literature..
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